The imprinting variance was estimated as the sum of both corresponding genetic variances per animal minus twice the covariance. The null hypothesis of no imprinting was tested by a restricted maximum likelihood ratio test with two degrees of freedom.
Genomic imprinting significantly contributed to the genetic variance of 19 traits. The proportion of the total additive genetic variance that could be attributed to genomic imprinting was of the order between 5% and 19%.
N. Neugebauer, H. Luther and N. Reinsch. Parent-of-origin effects cause genetic variation in pig performance traits. animal. 2010. Vol. 4 (5): 672-681.